RNDr. Václav Gvoždík Ph.D.

Position full-time researcher
Direction evolutionary history, herpetology

+420 560 590 622

Research interests

  • Evolution, speciation and phylogenetic diversification of vertebrates using molecular-genetic approaches, analyses of phenotype and biogeographic applications.
  • phylogeography, phylogenetics and biogeography of vertebrates of the Palearctic and Afrotropic ecozones,
  • molecular systematics and integrative taxonomy of amphibians and reptiles,
  • biodiversity of Africa and its conservation, Congo Basin,
  • herpetology.

Running Projects

Genetic networks in the Congolian forests: Unexplored biodiversity archive, herpetological aspect (2023–2025)

Finished projects

Genomic insights into the evolutionary history and contact zones of slow-worm lizards (Anguis) (2018–2020)
Amphibian species diversification across sky-island and lowland rainforests in a spatial and ecological context: genome-wide and continental transect (2015–2017)

Effective species conservation relies on a good knowledge of the biodiversity richness. This further helps us to better understand inter-organismal relationships in ecosystems and evolutionary processes. Africa is one of the least scientifically-explored regions of the World, particularly the Congo Basin. Knowledge of the biodiversity and distribution patterns in Africa therefore deserves detailed study.

Running Projects

ANGUIOMICS: Genomic insights into the evolutionary history and contact zones of slow-worm lizards (Anguis) (2018-2020)

Finished Projects

Amphibian species diversification across sky-island and lowland rainforests in a spatial and ecological context: genome-wide and continental transect (2015-2017)

Running projects

  • Speciation patterns in sky islands – Evolutionary history of montane amphibians of Africa (in cooperation with University of Basel, Switzerland).

Education

  • 2003-2010: Charles University in Prague, Faculty of Science, Department of Zoology
  • 2003: undergraduate study – MSc. thesis: „Interspecific and intraspecific variation of Hyla savignyi and Hyla arborea tree frogs” (supervisor: Jiří Moravec, National Museum).
  • 2010: Ph.D. study – thesis: „Phenotypic and molecular approaches in the systematics of the Palearctic and Neotropic tree frogs, Hyla and Osteocephalus (Amphibia: Hylidae)“ (supervisor: Jiří Moravec, National Museum; advisor: Petr Kotlík, IAPG AS CR).

Employment

  • 2005–2007: Charles University in Prague, Faculty of Science, Department of Zoology – research assistant,
  • 2005–2013: Institute of Animal Physiology and Genetics AS CR, Liběchov (2005-2009 postgrad; 2010-2013 postdoc),
  • 2012–2013: University of Basel, Biogeography Research Group, Switzerland – postdoctoral fellowship,
  • 2006–present: National Museum, Department of Zoology, Prague – scientific researcher,
  • 2014–present: Institute of Vertebrate Zoology, Academy of Sciences of the Czech Republic, Brno – scientific researcher.

Academic service

  • editor of Herpetology Notes,
  • committee member of the Czech Herpetological Society,
  • reviewer for scientific journals, e.g. African Journal of Herpetology, Amphibia – Reptilia, Animal Biology, Asian Herpetological Research, Biochemical Systematics and Ecology, Biological Journal of the Linnean Society, BMC Evolutionary Biology, Contributions to Zoology, Current Zoology, Herpetologica, Journal of Biogeography, Journal of Heredity, Revista de Biología Tropical, Zoologica Scripta, Zoological Science,
  • reviewer for Czech and foreign grant agencies,
  • assessor for IUCN African amphibian assessment.

Main international collaboration

  • University of Basel, Biogeography Research Group, Basel, Switzerland (S. P. Loader),
  • Royal Belgian Institute of Natural Sciences, Joint Experimental Molecular Unit, Brussels, Belgium (Z. T. Nagy),
  • Comenius University, Faculty of Science, Department of Zoology, Bratislava, Slovakia (P. Mikulíček, D. Jandzik),
  • short stays at research institutes: Great Britain – Natural History Museum, London; Spain – National Museum of Natural Sciences, Madrid; Germany – Natural History Museum, Berlin, Senckenberg Natural History Collections, Dresden; Italy – Trento Science Museum; Switzerland – Muséum d histoire naturelle, Geneva; Austria – Natural History Museum, Vienna; Israel – Galilee Biotechnology Center, Kiryat Shmona; Belgium – Royal Museum for Central Africa, Tervuren,

fieldwork in several countries of Europe, Middle East, Caucasus, North Africa and Sub-Saharan Africa: Cameroon, Republic of the Congo, Democratic Republic of the Congo, Tanzania.

Current PhD students

  • Daniel Jablonski – Phylogeography of slow worms (Anguis) in the Balkans (Comenius University, Faculty of Science, Department of Zoology, Bratislava, Slovakia; advisor).
  • Matej Dolinay – Species diversification of African amphibians in spatial and ecological context (Institute of Botany and Zoology, Faculty of Science, Masaryk University Brno; supervisor).

Former students

  • Petr Kůs – Genetic variation of horseshoe bats (Rhinolophus) in the Eastern Mediterranean (advisor).
  • Jan Šrámek – Genetic diversity of the Western Palearctic bent-winged bats (Miniopterus) (advisor).
  • Helena Šifrová – Genetic variation of slow worms (Anguis) in the Czech Republic and Slovakia.

Offered topics of Bachelor and Master thesis

please enquire by email

Publications

2024

Altmanová M, Doležálková-Kaštánková M, Jablonski D, Strachinis I, Vergilov V, Vacheva E, Iannucci A, Choleva L, Ráb P, Moravec J, Gvoždík V (2024). Karyotype stasis but species-specific repetitive DNA patterns in Anguis lizards (Squamata: Anguidae), in the evolutionary framework of Anguiformes. Zoological Journal of the Linnean Society, :.
Clements H, Do Linh San E, Hempson G, Linden B, Maritz B, Monadjem A, Reynolds C, Siebert F, Stevens N, Biggs R, De Vos A, Blanchard R, Child M, Esler K, Hamann M, Loft T, Reyers B, Selomane O, Skowno A, Tshoke T, Abdoulaye D, Aebischer T, Aguirre-Gutiérrez J, Alexander GJ, Ali AH, Allan DG, Amoako E, Angedakin S, Aruna E, Avenant NL, Badjedjea G, Bakayoko A, Bamba-Kaya A, Bates MF, Bates PJJ, Belmain SR, Bennitt E, Bradley J, Brewster CA, Brown MB, Brown M, Bryja J, Butynski TM, Carvalho F, Channing A, Chapman CA, Cohen C, Cords M, Cramer JD, Cronk N, Cunneyworth PMK, Dalerum F, Danquah E, Davies-Mostert HT, de Blocq AD, De Jong YA, Demos TC, Denys C, Djagoun CAMS, Doherty-Bone TM, Drouilly M, du Toit JT, Ehlers Smith YC, Eiseb SJ, Fashing PJ, Ferguson AW, Fernández-García JM, Finckh H, Fischer C, Gandiwa E, Gaubert P, Gaugris JY, Gibbs DJ, Gilchrist JS, Gil-Sánchez JM, Githitho AN, Goodman PS, Granjon L, Grobler JP, Gumbi BC, Gvoždík V (2024). The bii4africa dataset of faunal and floral population intactness estimates across Africa's major land uses. Scientific Data, 11:1205 191.
Gvoždík V, Knytl M, Zassi-Boulou A, Fornaini NR, Bergelová B (2024). Tetraploidy in the Boettger's dwarf clawed frog (Pipidae: Hymenochirus boettgeri) from the Congo indicates non-conspecificity with the captive population. Zoological Journal of the Linnean Society, 200: 1034-1047.
Chaney T, Pauwels OSG, Nagy ZT, Gvoždík V, Kusamba C, Badjedjea G, Masudi FM, Akuboy JB, Ernst R, Trape J, Chirio L, Conradie W, Keates C, Wallach V, Zassi-Boulou A, Vaughan ER, Greenbaum E (2024). Phylogenetics and integrative taxonomy of African water snakes (Squamata: Colubridae: Grayia). Herpetological Monographs, 38: 1-52.

2023

Badjedjea G, Masudi FM, Dudu Akaibe B, Gvoždík V (2023). Predation by the Large Green-eyed Frog, Cryptothylax greshoffii (Schilthuis, 1889), on Osorio’s Spiny Reed Frog, Afrixalus osorioi (Ferreira, 1906), in Sankuru Nature Reserve, Democratic Republic of the Congo. Herpetology notes, 16: 439-4411210 .
Fornaini NR, Bergelová B, Gvoždík V, Černohorská H, Krylov V, Kubíčková S, Fokam EB, Badjedjea G, Evans BJ, Knytl M (2023). Consequences of polyploidy and divergence as revealed by cytogenetic mapping of tandem repeats in African clawed frogs (Xenopus, Pipidae). European Journal of Wildlife Research, 69: 81.
Greenbaum E, Pauwels OSG, Gvoždík V, Vaughan ER, Chaney T, Buontempo M, Aristote MM, Muninga WM, Engelbrecht HM (2023). Systematics of the Thirteen-scaled Green Snake Philothamnus carinatus (Squamata: Colubridae), with the description of a cryptic new species from Central and East Africa. African Journal of Herpetology, 72: 119-144.
Gvoždík V, Nečas T, Jablonski D, Moriarty Lemmon E, Lemmon AR, Jandzik D, Moravec J (2023). Phylogenomics of Anguis and Pseudopus (Squamata, Anguidae) indicates Balkan-Apennine mitochondrial capture associated with the Messinian event. Molecular Phylogenetics and Evolution, 180: 107674.
Knytl M, Fornaini NR, Bergelová B, Gvoždík V, Černohorská H, Kubíčková S, Fokam EB, Evans BJ, Krylov V (2023). Divergent subgenome evolution in the allotetraploid frog Xenopus calcaratus. Gene, 851: 146974.
Sales-Oliveira V, Altmanová M, Gvoždík V, Kretschmer R, Ezaz T, Liehr T, Padutsch N, Badjedjea G, Utsonomia R, Tanomtong A, Cioffi M (2023). Cross-species chromosome painting and repetitive DNA mapping illuminate the karyotype evolution in true crocodiles (Crocodylidae). Chromosoma, 132: 289-303.

2022

Badjedjea G, Masudi FM, Akaibe BD, Gvoždík V (2022). Amphibians of Kokolopori: an introduction to the amphibian fauna of the Central Congolian Lowland Forests, Democratic Republic of the Congo. Amphibian & Reptile Conservation, 16: 35-70.
Jaynes KE, Myers EA, Gvoždík V, Blackburn DC, Portik DM, Greenbaum E, Jongsma GFM, Rödel M, Badjedjea G, Bamba-Kaya A, Baptista NL, Akuboy JB, Ernst R, Kouete MT, Kusamba C, Masudi FM, McLaughlin PJ, Nneji LM, Onadeko AB, Penner J, Pinto PV, Stuart BL, Tobi E, Zassi-Boulou A, Leaché AD, Fujita MK, Bell RC (2022). Giant Tree Frog diversification in West and Central Africa: Isolation by physical barriers, climate, and reproductive traits. Molecular Ecology, 31: 3979-3998.
Liedtke HC, Lyakurwa J, Lawson LP, Menegon M, Garrido-Priego M, Mariaux J, Ngalason W, Channing A, Owen NR, Bittencourt-Silva GB, Wilkinson M, Larson JG, Gvoždík V, Loader SP (2022). Thirty years of amphibian surveys in the Ukagurus Mountains of Tanzania reveal new species, yet others are in decline. African Journal of Herpetology, 71: 119-138.
Nečas T, Kielgast J, Nagy ZT, Kusamba Chifundera Z, Gvoždík V (2022). Systematic position of the Clicking Frog (Kassinula Laurent, 1940), the problem of chimeric sequences and the revised classification of the family Hyperoliidae. Molecular Phylogenetics and Evolution, 174: 107514.

2021

Benkovský N, Moravec J, Gvoždíková Javůrková V, Šifrová H, Gvoždík V, Jandzik D (2021). Phenotypic differentiation of the slow worm lizards (Squamata: Anguis) across their contact zone in Central Europe. PeerJ, 9: e12482.
Blackburn DC, Nielsen SV, Ghose S, Burger M, Gonwouo LN, Greenbaum E, Gvoždík V, Hirschfeld M, Kouete MT, Kusamba C, Lawson D, McLaughlin PJ, Zassi-Boulou A, Rödel M (2021). Phylogeny of African Long-Fingered Frogs (Arthroleptidae: Cardioglossa) reveals recent allopatric divergences in coloration. Ichthyology and Herpetology, 109: 728-742.
Deepak V, Maddock ST, Williams R, Nagy ZT, Conradie W, Rocha S, Harris DJ, Perera A, Gvoždík V, Doherty-Bone TM, Kamei RG, Menegon M, Labisko J, Morel C, Cooper N, Day JJ, Gower D (2021). Molecular phylogenetics of sub-Saharan African natricine snakes, and the biogeographic origins of the Seychelles endemic Lycognathophis seychellensis. Molecular Phylogenetics and Evolution, 161: 107152.
Dolinay M, Nečas T, Zimkus BM, Schmitz A, Fokam EB, Lemmon EM, Lemmon AR, Gvoždík V (2021). Gene flow in phylogenomics: Sequence capture resolves species limits and biogeography of Afromontane forest endemic frogs from the Cameroon Highlands. Molecular Phylogenetics and Evolution, 163: 107258.
Gvoždík V, Harca Z, Hánová A, Jablonski D, Pupins M, Čeirāns A, Paasikunnas T (2021). Two species of slow worm (Anguis fragilis, A. colchica) present in the Baltic region. Amphibia-Reptilia, 42: 383-389.
Jablonski D, Sillero N, Oskyrko O, Bellati A, Čeirāns A, Cheylan M, Cogalniceanu D, Crnobrnja-Isailović J, Crochet P, Crottini A, Doronin I, Džukić G, Geniez P, Ilgaz Ç, Iosif R, Jandzik D, Jelić D, Litvinchuk S, Ljubisavljević K, Lymberakis P, Mikulíček P, Mizsei E, Moravec J, Najbar B, Pabijan M, Pupins M, Sourrouille P, Strachinis I, Szabolcs A, Thanou E, Tzoras E, Vergilov V, Vörös J, Gvoždík V (2021). The distribution and biogeography of slow worms (Anguis, Squamata) across the Western Palearctic, with an emphasis on secondary contact zones. Amphibia-Reptilia, 42: 519-530.
Lokasola A, Lotana C, Tungaluna G, Akaibe BD, Jirků M, Gvoždík V (2021). New distributional data and genetic variation of Panaspis breviceps (Squamata: Scincidae) indicate a biogeographic connection across the Congo Basin. African Zoology, 56: 312-318.
Nečas T, Badjedjea G, Vopálenský M, Gvoždík V (2021). Congolius, a new genus of African reed frog endemic to the central Congo: A potential case of convergent evolution. Scientific Reports, 11: 8338.

2020

Gvoždík V, Nečas T, Dolinay M, Zimkus BM, Schmitz A, Fokam EB (2020). Evolutionary history of the Cameroon radiation of puddle frogs (Phrynobatrachidae: Phrynobatrachus), with descriptions of two critically endangered new species from the northern Cameroon Volcanic Line. PeerJ, 8: e8393.
Vlček P, Zavadil V, Gvoždík V (2020). The need for transboundary faunistics and conservation: first record of the Natterjack Toad (Epidalea calamita) in Czech Silesia, northeastern Czech Republic. Amphibian & Reptile Conservation, 14: 62-69 e254.
Zimkus BM, Baláž V, Belasen AM, Bell RC, Channing A, Doumbia J, Fokam EB, Gonwouo LN, Greenbaum E, Gvoždík V, Hirschfeld M, Jackson K, James TY, Kusamba C, Larson JG, Mavoungou L, Rödel M, Zassi-Boulou A, Penner J (2020). Chytrid pathogen (Batrachochytrium dendrobatidis) in African amphibians: a continental analysis of occurrences and modeling of its potential distribution. Herpetologica, 76: 201-215.

2019

Babangenge GB, Jocqué R, Masudi FM, Rödel M, Burger M, Gvoždík V, Pauwels OSG (2019). Frog-eating spiders in the Afrotropics: an analysis of published and new cases. Bulletin of the Chicago Herpetological Society, 54: 57-631210 .
Evans BJ, Gansauge MT, Stanley EL, Furman BLS, Cauret CMS, Ofori-Boateng C, Gvoždík V, Streicher J, Greenbaum E, Tinsley RC, Meyer M, Blackburn DC (2019). Xenopus fraseri: Mr. Fraser, where did your frog come from?. PLoS ONE, 14: e0220892.
Ihlow F, Farooq HM, Gvoždík V, Hofmeyr MD, Conradie W, Campbell PD, Harvey J, Verburgt L, Fritz U (2019). Geographic range extension of Speke’s Hinge-back Tortoise Kinixys spekii Gray, 1863. Amphibian & Reptile Conservation, 13: 61-67 e195.
Jablonski D, Gvoždík V, Choleva L, Jandzik D, Moravec J, Mačát Z, Veselý M (2019). Tracing the maternal origin of the common wall lizard (Podarcis muralis) on the northern range margin in Central Europe. Mitochondrion, 46: 149-157.
Leaché AD, Portik DM, Rivera D, Rödel M, Penner J, Gvoždík V, Greenbaum E, Jongsma GFM, Ofori-Boateng C, Burger M, Eniang EA, Bell RC, Fujita MK (2019). Exploring rain forest diversification using demographic model testing in the African foam-nest treefrog Chiromantis rufescens. Journal of Biogeography, 46: 2706-2721.
Portik DM, Bell RC, Blackburn DC, Bauer AM, Barratt CD, Branch WR, Burger M, Channing A, Colston TJ, Conradie W, Dehling JM, Drewes RC, Ernst R, Greenbaum E, Gvoždík V, Harvey J, Hillers A, Hirschfeld M, Jongsma GFM, Kielgast J, Kouete MT, Lawson LP, Leaché AD, Loader SP, Lötters S, Meijden A, Menegon M, Müller S, Nagy ZT, Ofori-Boateng C, Ohler A, Papenfuss TJ, Rößller D, Sinsch U, Rödel M, Veith M, Vindum J, Zassi-Boulou A, McGuire JA (2019). Sexual dichromatism drives diversification within a major radiation of African amphibians. Systematic Biology, 68: 859-875.

2018

Dolinay M, Taboue GCT, Fokam EB, Gvoždík V (2018). Geographic and altitudinal range extension of Arthroleptis palava (Anura: Arthroleptidae) from the Cameroon Volcanic Line. Herpetology notes, 11: 557-5601210 .
Gvoždík V, Dolinay M, Zassi-Boulou A, Babangenge GB (2018). New data on Trachylepis makolowodei from Central Africa. Herpetology notes, 11: 515-5181210 .
Charles KL, Bell RC, Blackburn DC, Burger M, Fujita MK, Gvoždík V, Jongsma GFM, Kouete MT, Leaché AD, Portik DM (2018). Sky, sea, and forest islands: Diversification in the African leaf-folding frog Afrixalus paradorsalis (Anura: Hyperoliidae) of the Lower Guineo-Congolian rain forest. Journal of Biogeography, 45: 1781-1794.
Jandzik D, Jablonski D, Zinenko O, Kukushkin OV, Moravec J, Gvoždík V (2018). Pleistocene extinctions and recent expansions in an anguid lizard of the genus Pseudopus. Zoologica Scripta, 47: 21-32.
Kotsakiozi P, Jablonski D, Ilgaz Ç, Kumlutaş Y, Avci A, Meiri S, Itescu Y, Kukushkin O, Gvoždík V, Scillitani G, Roussos SA, Jandzik D, Kasapidis P, Lymberakis P, Poulakakis N (2018). Multilocus phylogeny and coalescent species delimitation in Kotschy's gecko, Mediodactylus kotschyi: Hidden diversity and cryptic species. Molecular Phylogenetics and Evolution, 125: 177-187.
Mikulíček P, Jablonski D, Páleník M, Gvoždík V, Jandzik D (2018). Characterization of microsatellite markers in the genera Anguis and Pseudopus (Reptilia: Anguidae). Salamandra, 54: 158-1621210 .

2017

Bell RC, Parra JL, Badjedjea G, Barej MF, Blackburn DC, Burger M, Channing A, Dehling JM, Greenbaum E, Gvoždík V, Kielgast J, Kusamba C, Lötters S, McLaughlin PJ, Nagy ZT, Rödel M, Portik DM, Stuart BL, VanDerWal J, Zassi-Boulou A, Zamudio KR (2017). Idiosyncratic responses to climate-driven forest fragmentation and marine incursions in reed frogs from Central Africa and the Gulf of Guinea Islands. Molecular Ecology, 26: 5223-5244.
Doherty-Bone TM, Gvoždík V (2017). The amphibians of Mount Oku, Cameroon: an updated species inventory and conservation review. ZooKeys, 643: 109-139.
Jablonski D, Najbar B, Grochowalska R, Gvoždík V, Strzała T (2017). Phylogeography and postglacial colonization of Central Europe by Anguis fragilis and Anguis colchica. Amphibia-Reptilia, 38: 562-569.
Lehr E, Moravec J, Cusi JC, Gvoždík V (2017). A new minute species of Pristimantis (Amphibia: Anura: Craugastoridae) with a large head from the Yanachaga-Chemillén National Park in central Peru, with comments on the phylogenetic diversity of Pristimantis occurring in the Cordillera Yanachaga. European Journal of Taxonomy, 325: 1-221210 .
Zimkus BM, Lawson LP, Barej MF, Barratt CD, Channing A, Dash KM, Dehling JM, Du Preez L, Gehring P, Greenbaum E, Gvoždík V, Harvey J, Kielgast J, Kusamba C, Nagy ZT, Pabijan M, Penner J, Rödel M, Vences M, Lötters S (2017). Leapfrogging into new territory: How Mascarene ridged frogs diversified across Africa and Madagascar to maintain their ecological niche. Molecular Phylogenetics and Evolution, 106: 254-269.

2016

Jablonski D, Jandzik D, Mikulíček P, Džukić G, Ljubisavljević K, Tzankov N, Jelić D, Thanou E, Moravec J, Gvoždík V (2016). Contrasting evolutionary histories of the legless lizards slow worms (Anguis) shaped by the topography of the Balkan Peninsula. BMC Evolutionary Biology, 16: 99.
Liedtke HC, Müller H, Rödel M, Menegon M, Gonwouo LN, Barej MF, Gvoždík V, Schmitz A, Channing A, Nagel P, Loader SP (2016). No ecological opportunity signal on a continental scale? Diversification and life-history evolution of African true toads (Anura: Bufonidae). Evolution, 70: 1717-1733.

2015

Bell RC, Drewes RC, Channing A, Gvoždík V, Kielgast J, Lötters S, Stuart BL, Zamudio KR (2015). Overseas dispersal of Hyperolius reed frogs from Central Africa to the oceanic islands of Sao Tomé and Príncipe. Journal of Biogeography, 42: 65-75.
Evans BJ, Carter TF, Greenbaum E, Gvoždík V, Kelley DB, McLaughlin PJ, Pauwels OSG, Portik DM, Stanley EL, Tinsley RC, Tobias ML, Blackburn DC (2015). Genetics, morphology, advertisement calls, and historical records distinguish six new polyploid species of African clawed frog (Xenopus, Pipidae) from West and Central Africa. PLoS ONE, 10: e0142823.
Furman BLS, Bewick AJ, Harrison TL, Greenbaum E, Gvoždík V, Kusamba C, Evans BJ (2015). Pan-African phylogeography of a model organism, the African clawed frog "Xenopus laevis". Molecular Ecology, 24: 909-925.
Gvoždík V, Canestrelli D, García-París M, Moravec J, Nascetti G, Recuero E, Teixeira J, Kotlík P (2015). Speciation history and widespread introgression in the European short-call tree frogs (Hyla arborea sensu lato, H. intermedia and H. sarda). Molecular Phylogenetics and Evolution, 83: 143-155.
Penske S, Gvoždík V, Menegon M, Loader SP, Müller H (2015). Description of the tadpole of Leptopelis cf. grandiceps (Amphibia: Anura: Arthroleptidae) from the Uluguru Mountains, Tanzania. Herpetological Journal, 25: 61-64.

2014

Barej MF, Rödel M, Loader SP, Menegon M, Gonwouo N, Penner J, Gvoždík V, Günther R, Bell RC, Nagel B, Schmitz A (2014). Light shines through the spindrift – Phylogeny of African torrent frogs (Amphibia, Anura, Petropedetidae). Molecular Phylogenetics and Evolution, 71: 261-273.
Gvoždík V, Tillack F, Menegon M, Loader SP (2014). The status of Leptopelis barbouri Ahl, 1929 and eleven other nomina of the current tree-frog genus Leptopelis (Arthroleptidae) described from East Africa, with a redescription of Leptopelis grandiceps Ahl, 1929. Zootaxa, 3793: 165-187.
Nagy ZT, Gvoždík V, Meirte D, Collet M, Pauwels OSG (2014). New data on the morphology and distribution of the enigmatic Schouteden's sun snake, Helophis schoutedeni (de Witte, 1922) from the Congo Basin. Zootaxa, 3755: 96-100.

2013

Nagy ZT, Kusamba C, Collet M, Gvoždík V (2013). Notes on the herpetofauna of western Bas-Congo, Democratic Republic of the Congo. Herpetology notes, 6: 413-4191210 .